core histones linker DNA of nucleosome “beads-on-a-string nucleosome includes form of ch pairs of DNA NUCLEASE DIGESTS LINKER DNA released nucleosome 11 nm core particle DISSOCIATION WITH HIGH CONCENTRATION OF SALT 05 octameric 146-nucleotide-pair histone core DNA double helix DISSOCIATION ⑦6 ⑦④， 9 H2A H2B H3 H4