PEP cytosolic PEP carboxykinase Oxaloacetate NADH+H' cytOsole malate Malate Malate PEP NADT mitochondrial PEP NADH+H carboxylase Oxaloacetate Oxaloacetate carboxylase pyruvate carboxylase Co Pyruvate Pyruvate Mitochondrion novate Pyruvate NADH+H+ iaeste NAD Lactate丙 酮 酸 生 成 磷 酸 烯 醇 式 丙 酮 酸