activities in soil -the drilosphere (After Beare et al, 1995) feces 的? + casts detritusphere N20 02c02 uncus soil -the porosphere water, (After Beare et al, 1995 nitr ate H20 eaf N03 DOC leaf litter :nutrient translocation hy pha mineral so N2 aggregatusphere N20 C02 rhizosphere CovA s and bacteriamILno c humus hypha NH4 ANHA amoeba N03 organIc atter