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Acknowledgements This work was supported by the EPSRC (grant numbers EP/M009580/1, EP/K016288/1 and EP/L016354/1), the Royal Society and the Leverhulme Trust. O.I. acknowledges support from DOD-ONR (N00014-16- 1-2311) and an Eshelman Institute for Innovation award. Reviewer information Nature thanks F.-X. Coudert, M. Waller and the other anonymous reviewer(s) for their contribution to the peer review of this work. Author contributions All authors contributed equally to the design, writing and editing of the manuscript. Competing interests The authors declare no competing interests. Additional information Reprints and permissions information is available at http://www.nature.com/ reprints. Correspondence and requests for materials should be addressed to O.I. or A.W. Publisher’s note: Springer Nature remains neutral with regard to jurisdictional claims in published maps and institutional affiliations. 26 J U LY 2018 | V OL 559 | NATUR E | 555 © 2018 Springer Nature Limited. 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