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presumptive RPE Noggin lens pit TGFB Pax6+/Sox1+ E8-E9 SE Shh。 ↓ presumptive neural retina Six3+ optic stalk Rx+ Rx E9.5 ↓ ↓ Crx+/Chx10+ Otx2+/Mitf+ Rhodopsin RPE65+ E10 NR RPERx+ NR Noggin Six3+ Crx+/Chx10+ Rhodopsin RPE RPE65+ Otx2+/Mitf+ RxPax6+/Sox1+