口一个同胞k次度量表型值的均值P或Ps k k 0.5h 0.25h 1+(k-1) HS 1+(k-1)r k b=0.5h2 k 0.25h 1+(k-1) 1+(k-1) 0.5kh2 0.25kh2 1+(k-1)l FS 1s1+(k-1)r2S 一个同胞 次度量表型值的均值 或            + − = + − = + − = H S e H S e AP e AP P k r k h A k r k b h k r k r h H S H S 1 ( 1) 0.25 ˆ 1 ( 1) 0.25 1 ( 1) 0.25 2 2            + − = + − = + − = FS e FS e AP e AP P k r k h A k r k b h k r k r h FS FS 1 ( 1) 0.5 ˆ 1 ( 1) 0.5 1 ( 1) 0.5 2 2 k PFS PHS