Nutrient cycles in forest e cosyste ms Atmospheric inputs Tra nsloc a tion l一Foliar uptake Foliage Foliar Leaching Fertilization,Recycling Ltter Fa / Forest Floor Harvest Root up缸ke Hum us Mine ra za tio n Litter Leaching Mine raliza tion Soil We athering Soil Unavaila ble Ava ilable Pools Pools Im mobilzation Adsorp tion Soil Le aching Output Atmospheric inputs Output FoliageWo o d Fore st Floor Soil Availa ble Pools Soil Unavaila ble Pools Tra nsloc a tion Foliar Lea ching Litter Fall Foliar uptake Root up ta ke Hum us Mine ra liza tion Litter Leac hing Mine ralization We athering Im mobilization Adsorp tion Nutrient c yc les in forest ec osystems Soil Le ac hing Harvest Fer4liza4on, Recycling