The Nobel Prize in Physiology or Medicine 1968 for their interpretation of the genetic code and its function in protein synthesis" Presentation Speech Established the chemical structure of tRNA Robert W.Holley USA Cornell University Ithaca,NY,USA 1922-1993 Biography Devised methods to synthesize Established the in vitro system well defined nucleic acids for revealing the genetic codes Har Gobind Marshall W. Khorana Nirenberg USA USA University of National Institutes Wisconsin of Health Madison,WI,USA Bethesda,MD.USA 1922 1927 Established the chemical structure of tRNA Established the in vitro system for revealing the genetic codes Devised methods to synthesize well defined nucleic acids