1958 Joshua Lederberg ( Phage transduction Beadle&Tatum(美) One gene-one enzyme 红色面包霉突变体 Joshua Lederberg Edward George Wells Lawrie Tatum Beadle 1958 Joshua Lederberg（美） Phage transduction Beadle & Tatum（美） One gene-one enzyme 红色面包霉突变体 George Wells Beadle Edward Lawrie Tatum Joshua Lederberg