Hydrolysis of Complex Carbohydrates (b)Inverting glycosidases operate through a single inversion by nucleophilic attack of water through an SN2-like inversion (b)Inverting glycosidase Enz CH2OH CH2OH HO HO OH OH OH Oxonium ion CH2OH HO -0 HO Cengage Learning.All Rights Reserved.(b) Inverting glycosidases operate through a single inversion by nucleophilic attack of water through an SN2-like inversion Hydrolysis of Complex Carbohydrates