△C ATP丿S ATP Cy·△t+ maX △Cx X/ATP (△CAmp)s=m`Cy+ △Cx △t maX △t X/ATP (△CAP)s=m/TP △C X Cy·▲ max Cy·△t X/ATP qATP matp t max X/ATP则 ( ) ( ) ( ) = +      = +       =  +     =    +   max / max / max / max / 1 1 1 1 X ATP ATP ATP X X X ATP ATP X ATP S X X ATP ATP X ATP S X X ATP ATP S ATP X Y q m C t C Y m C t C t C Y m C t C C Y C m C t