Factor Compensation Compensation along gradients of temperature. light,pH,or other factors usually involves genetic changes of ecotypes,but it can occur by physiological adjustments without genetic fixation. Ca and Sr锶（锶在一定条件下，软体动物贝壳用 锶替代部分钙) Factor Compensation Factor Compensation • Compensation along gradients of temperature, light, pH, or other factors usually involves genetic changes of ecotypes, but it can occur by physiological adjustments without genetic fixation. • Ca and Sr 锶 (锶在一定条件下,软体动物贝壳用 锶替代部分钙 )