Some subsidiary补充principles 1.Organisms may have a wide range of tolerance for one factor and a narrow range for another. 2.Organisms with wide ranges of tolerance for limiting factors are likely to be most widely distributed. 3.When conditions are not optimal for a species with respect to one ecological factor,the limits of tolerance may be reduced for other ecological factors. Some subsidiary Some subsidiary 补充 principles principles 1. Organisms may have a wide range of tolerance for one factor and a narrow range for another. 2. Organisms with wide ranges of tolerance for limiting factors are likely to be most widely distributed. 3. When conditions are not optimal for a species with respect to one ecological factor, the limits of tolerance may be reduced for other ecological factors