化学渗透假说 Outer membrane Inter- membrane 4 HO >4H 2 H space ④t ODORa Inner Cyto- mitochon 从‖戏人k drial 4 H 2HO 4 Hb wHO 1202c2623 Matrix space 2-3 2-3 aePP Citric acid ATP NADH cycle H2O 10H B-Oxidation B Organization化学渗透假说