○葡调目的常见科和李程至要科的特征。 目的 部兴器器益,w平 一、分科特行 以及青餐红损式。复限形状和省生位五。触角后状,外烟片的有无,前房有板的影软。前足基性窝类型,足的类型、图世数和形软。首后超类型。 二。分科检案表 豹翅目分科检素表 1前胸背侧维明显:腹部第1节腹板核后足基节窝完全分开 后足基节达髓边:第1板节不可见:触角多光:水生 前两背板比头窄:触角间距小于上唇的宽 前足最短,后足最长:复限正常】 不 Dde 星不长外雄2条:有外聘片 三锥象甲科Brentidae 7 .10 11附节隐5节 附节非隐5节 12 豆象科Bruchidae 天牛科Crabid 育面降起:老数种类有后基纷 .16 16前胸腹板有1个限形突 .17 前腹板无形实 18 6 1 19翰翻柔软 .20 ·鹬翅坚硬 20有发光:前背板多为圆形头前背背长住. .21 -22 触角非鰓片状 27 ,锹甲科Lucanidae 24 24后足径节1端距:小盾片常不可见:中足基节运离:触角8一9竹. ,金鱼甲科Scarabacidac
目 的 认识鞘翅目的常见科和掌握重要科的特征。 材 料 虎甲,步甲,龙虱,豉甲,水龟虫,隐翅虫,萤火虫(♀♂),鳃金龟,丽金龟,花金龟,独角仙(♀♂),锹甲(♀♂),叩头虫,吉丁虫,天 牛,叶甲,拟步甲,芫青,瓢虫,豆象,象甲,三锥象甲,小蠹虫,幼虫类型,以及本次实验各科示范标本。 内容与方法 一、分科特征 主要包括头式,复眼形状和着生位置,触角形状,外咽片的有无,前胸背板的形状,前足基节窝类型,足的类型、跗节数和形状,前后翅类型, 以及腹部节数等。 二、分科检索表 鞘翅目分科检索表 1 前胸背侧缝明显;腹部第1节腹板被后足基节窝完全分开················································································ 2 - 前胸很少有背侧缝;腹部第1节腹板不被后足基节窝完全分开········································································· 5 2 后足基节不达鞘翅边缘;第1腹节可见;触角丝状具毛;陆生········································································ 3 - 后足基节达鞘翅边缘;第1腹节不可见;触角多光滑;水生············································································· 4 3 前胸背板比头宽;触角间距大于上唇的宽·················································································· 步甲科Carabidae - 前胸背板比头窄;触角间距小于上唇的宽··············································································· 虎甲科Cicindelidae 4 前足长,中后足明显短于前足;每侧复眼分上下两部分························································· 豉甲科Gyrinidae - 前足最短,后足最长;复眼正常,不分成两部分····································································· 龙虱科Dytiscidae 5 头部延长呈喙状;外咽缝l条;无外咽片············································································································· 6 - 头部正常,不延长;外咽缝2条;有外咽片········································································································· 8 6 喙长又直,触角丝状·············································································································· 三锥象甲科Brentidae - 喙短,若较长即触角膝状 ········································································································································· 7 7 喙长又明显;前胸背板不包住头部;胫节外侧无齿列,端部也无弯距··························· 象甲科Curculionidae - 喙短,不明显;前胸背板包住头部一部分;胫节外侧具齿列或端部有弯距························· 小蠹科Scolytidae 8 下颚须长于或等于触角;中胸腹面有1个大纵刺·························································· 水龟虫科Hydrophilidae - 下颚须短于触角;中胸腹面无纵刺························································································································· 9 9 前足和中足跗节5节,后足跗节4节·················································································································· 10 - 各足跗节数相等 ······················································································································································· 11 10 前足基节窝后方封闭;爪不分裂······················································································· 拟步甲科Tenebrionidae - 前足基节窝后方开放;爪分裂为2叉状······················································································· 芫青科Meloidae 11 跗节隐5节 ······························································································································································ 12 跗节非隐5节 ·························································································································································· 14 12 头部短喙状;鞘翅短,腹末外露;前胸背板梯形;后足基节左右靠近································· 豆象科Bruchidae - 头部不呈喙状;鞘翅长,腹末多不外露;前胸背板长形;后足基节左右分开··············································· 13 13 复眼肾形,环绕触角;触角长于体长的一半······································································· 天牛科Cerambycidae - 复眼圆形,不环绕触角;触角短于体长的一半·································································· 叶甲科Chrysomelidae 14 鞘翅短,末端平截;腹末露出大部分··············································································· 隐翅虫科Staphylinidae - 鞘翅普通长形,末端不平截,盖住腹部大部分··································································································· 15 15 跗节隐4节;体半球形,背面隆起;多数种类有后基线···················································· 瓢虫科Coccinellidae - 跗节式5-5-5;无后基线 ········································································································································· 16 16 前胸腹板有1个楔形突 ·········································································································································· 17 - 前胸腹板无楔形突 ··················································································································································· 18 17 前胸背板与中胸连接紧密,不能动;前胸背板与鞘翅相接处不凹下······························ 吉丁虫科Buprestidae - 前胸背板与中胸连接不紧密,可动;前胸背板与鞘翅相接处凹下······································ 叩头虫科Elateridae 18 多有1个中单眼;后足基节有容纳腿节的槽;足极短,常缩于体下································· 皮蠹科Dermestidae - 无中单眼;后足基节无容纳腿节的槽,若有此槽,则足不短··········································································· 19 19 鞘翅柔软 ·································································································································································· 20 - 鞘翅坚硬 ··································································································································································· 21 20 有发光器;前胸背板多为半圆形;头常被前胸背板盖住·························································· 萤科Lampyridae - 无发光器;前胸背板多为方形;头不被前胸背板盖住··························································· 花萤科Cantharidae 21 触角鳃片状 ······························································································································································ 22 - 触角非鳃片状 ··························································································································································· 27 22 触角鳃片部各节不呈叶形,不能相互并合;雄虫上颚发达如鹿角状····································· 锹甲科Lucanidae - 触角鳃片部各节呈扁平叶形,能相互并合;雄虫上颚正常··············································································· 23 23 腹部气门全部为鞘翅所覆盖·································································································································· 24 - 腹部气门不全部为鞘翅所覆盖······························································································································· 25 24后足胫节1端距;小盾片常不可见;中足基节远离;触角8~9节································ 金龟甲科Scarabaeidae
26 ,花金龟科Cetoniidae 下口式赠节式55-5.幼虫式,第5腹节背面突起上有逆钩。常见种类中华虎甲Cicindel chinensis DeGeer 养养 图42步甲科 A.虎甲科:B步甲科:C龙科 运离前中足:虫前足抱程足:胎节式55。幼虫式。服末有尾突,常见种类:黄边龙 这离中后足:节式555。幼虫式,腹竹两侧有气管鳃,端都有尾钩。常见种 类:大政甲Dineulus orientalis moodee 0意营 图43鞘翅日分科 一B.破甲科:CD水龟甲科 (二)多食亚Polypha 体水大电细说玻得》长于支等于装角长:头有外系片:多质种美中质面有们个黄关:后节式5,衣是特类:长美水心 独角10节:头部有外用片:粉常极短,端平:后翅折叠于翻翅之下:腹节器出3节以上:附节式5-5-5,2-2-2或3-3-3.常见种类:青蚁 手海恶
- 后足胫节2端距;小盾片发达;中足基节十分接近;触角11节···································· 粪金龟科Geotrupidae 25 腹部气门多位于腹板侧端;每侧气门几乎成直线排列;色彩暗淡······························ 鳃金龟科Melolonthidae - 腹部前部数对气门常位于侧膜;每侧气门成折线排列;色彩艳丽,有金属光泽··········································· 26 26后足端跗节2爪长度不等,1爪末端常分叉;鞘翅缘折于肩后不内弯································ 丽金龟科Rutelidae - 后足端跗节2爪长度相等,末端不分叉;鞘翅缘折于肩后向内弯····································· 花金龟科Cetoniidae 27 体形似蜘蛛;触角线状或念珠状;前胸背板前缘不凹入;第1跗节正常································ 蛛甲科Ptinidae - 体形正常;触角端1~3节膨大;前胸背板前缘凹入;第1跗节细小······························ 谷盗科Trogossitidae 三、常见科的主要特征和代表种类 (一)肉食亚目Adephaga 1.虎甲科Cicindelidae(图42A) 触角11节;头部有外咽片,常宽于前胸,下口式;跗节式5-5-5。幼虫蛃式,第5腹节背面突起上有逆钩。常见种类:中华虎甲Cicindela chinensis DeGeer。 2.步甲科Carabidae(图42B) 触角11节;头部有外咽片,常狭于前胸,前口式;跗节式5-5-5。幼虫蛃式,第9腹节有伪足状突起。常见种类:中华步甲Carabus maderae chinensis Kirby。 图42 步甲科 A.虎甲科;B.步甲科;C.龙虱科 3.龙虱科Dytiscidae(图42C) 触角11节;体背弧形;头部有外咽片;后足最长,远离前中足;雄虫前足抱握足;跗节式5-5-5。幼虫蛃式,腹末有尾突。常见种类:黄边龙虱 Cybister japonicus Sharp。 4.豉甲科Gyrinidae(图43A,B) 触角粗短;头部有外咽片;复眼分为上下两个;前足最长,远离中后足;跗节式5-5-5。幼虫蛃式,腹节两侧有气管鳃,端部有尾钩。常见种 类:大豉甲Dineulus orientalis Moodeer。 图43 鞘翅目分科 A~B.豉甲科;C~D.水龟甲科 (二)多食亚目Polyphaga 5.水龟甲科Hydrophilidae(图43C,D) 体背弧形;触角短棍棒状;下颚须丝状,长于或等于触角长;头部有外咽片;多数种类中胸腹面有1个刺突;跗节式5-5-5。常见种类:长须水龟 甲Hydrophilus acuminatus Motsch.。 6.隐翅虫科Staphylinidae(图44A) 触角10节;头部有外咽片;鞘翅常极短,末端平截;后翅折叠于鞘翅之下;腹节露出3节以上;跗节式5-5-5,2-2-2或3-3-3。常见种类:青翅蚁 形隐翅虫Paederus fuscipes Curtis
人隐是虫,BC金电型是是分”E花金他新:E保全电新 股角计:背调背发益住头部:葡题柔软:留节式55。华虫常无超,发光器位于腹木装1节腹面。性虫有超,发光器他于腹来2节腹面。薇见 个三系我标于用后不内家中后足收器前中之大分在 a (Bates). 角形:翅缘折于肩后向内:中后足有2端距:前中足的爪不分又。常见种类:雅静花金鱼C0m10m0rpb 触角状,10节,片部3一6节:头大,前口式,上颗发达,尤以虫明显:前青板很宽,方形:可见腹板5个。大型种类:福运甲 前胸背板与稍翅相接处同在同一弧线上:前病与中胸密接,不可动:附节式555:幼虫无足型,前胸背板两面呈盾状,宽于头部
图44 鞘翅目分科 A.隐翅虫科;B~C.金龟科;D.鳃金龟科;E.花金龟科;F.犀金龟科 7.萤科Lampyridae 触角11节;前胸背板盖住头部;鞘翅柔软;跗节式5-5-5。雌虫常无翅,发光器位于腹末端1节腹面;雄虫有翅,发光器位于腹末2节腹面。常见 种类:中华黄萤Luciola chinensis L.。 8.金龟甲科Scarabaeidae(图44B,C) 触角8~9节,鳃片部3节;中胸腹板侧片达足之基节,小盾片不可见;后足胫节有1枚端距;腹部有6个可见腹板;跗节式5-5-5。著名种类:神农 洁蜣螂Catharsius molossus(L.)。 9.粪金龟科Geotrupidae 触角11节,鳃片部3节;前足拟开掘足;中胸小盾片发达;鞘翅常有纵沟线;后足胫节有2枚端距;有6个可见腹板。常见种类:戴锤角粪金龟 Bolbotrypes davidis Fairmaire。 10.丽金龟科Rutelidae 触角多9节,少数10节,鳃片部3节;小盾片三角形;鞘翅缘折于肩后不内弯;中后足有2枚端距;前中足2爪之大爪常分叉。常见种类:铜绿异 丽金龟Anomala corpulenta Mots.。 11.鳃金龟科Melolonthidae(图44D) 触角8~10节,鳃片部3~8节,以3节最多;小盾片三角形;前胸背板通常宽大于长,基部等于或窄于鞘翅基部;鞘翅常有4条纵肋可见;中后足 有2枚端距。常见种类:东北大黑鳃金龟Holotrichia diomphalia (Bates)。 12.花金龟科Cetoniidae(图44E) 触角10节,鳃片部3节;小盾片三角形;鞘翅缘折于肩后向内弯;中后足有2枚端距;前中足的爪不分叉。常见种类:雅静花金龟Cosmiomorpha modesta Saunders。 13.犀金龟科Dynastidae(图44F) 触角10节,鳃片部3节;头和前胸背板有角突,在雄虫中尤其显著;前胸腹板有柱形、三角形、舌形等突起;小盾片三角形。常见种类:双叉犀 金龟Allomyrina dichotoma (L.)。 14.锹甲科Lucanidae(图45C) 触角膝状,10节,鳃片部3~6节;头大,前口式,上颚发达,尤以雄虫明显;前胸背板很宽,方形;可见腹板5个。大型种类:福运锹甲 Lucanus fortunei Saunders。 图45 鞘翅目分科 A.锹甲科;B.叩甲科;C.吉丁虫科科 15.叩甲科Elateridae(图45B,46A,B) 触角11~12节;前胸背板与鞘翅相接处凹下;前胸腹板有1个楔突插入中胸腹板沟内,作为弹跳工具;跗节式5-5-5。幼虫蛃型,表皮黄褐色且坚 硬,又称金针虫。重要种类:沟叩甲Pleonomus canaliculatus (Faldermann)。 16.吉丁虫科Buprestidae(图45C) 触角11节;前胸背板与鞘翅相接处同在同一弧线上;前胸与中胸密接,不可动;跗节式5-5-5;幼虫无足型,前胸背板两面呈盾状,宽于头部。 常见种类:柑橘吉丁甲Agrilus auriventris Saunders。 17.负泥虫科Crioceridae 触角11节,丝状、棒状、锯齿状或栉状;前胸背板长大于宽;足胫节端部常有距;后足腿节粗大,内侧具齿,胫节弯曲;腹部腹面可见5节,部 分种类第1可见腹节很长,为其余各节长度之和。幼虫头小,胸部各节较腹部小,无腹足及尾突。常见种类:绿豆象Callosobruchus chinensis (L.)。 18.天牛科Cerambycidae(图46C~D) 触角丝状,11节,长于体长之半;复眼内缘内凹;跗节隐5节。幼虫无足型,头部多缩入前胸内,胸腹背面多隆突。重要种类:松褐天牛 Monochamus alternatus Hope
sn前胸腹板:s1m2中胸腹板 sn后胸夜板:cX1前足基节 cx2中足基cx3后足基节 sp前胸腹板刺:cx基节 少数 0节 不及休长之半:复眼哪圆 附节 5节 幼虫型。重要种美:黄曲条跳 Phyllotreta (Fabricius). 人头:B时D:E出 影极生甲新猫行发记球折:前足装节实闲大:雨节式554。幼虫以金针血。自腹木无金针虫那行的骨质突起和伪足。重要问 头与前胸等宽:下口式:翅较软:前足基节窝开放式:附节式554:爪二分裂。幼虫复变态,食蝗虫卵等。药用种类:大斑青yb ata (Pal 电有护”限然品蛋:男隐。多为食,少为食内食性成体有无毛,有光泽,总角省生于丙复 触角9一11节。非膝状:嚎细长,直,约与前胸等长:前足基节半圆形:后腹板约与腹部等长:可见腹板5节:节隐5节。常见种类:甘小 象甲Cylas/ormicarius(Fabricius) 米《佩 图49鞘日分利 A.三锥象甲科:B.象甲科C.小鳌科 站鲁现g时得园市.本科为的是是大.重要群表:音季p加noL小
图46 鞘翅目分科 A~B.叩甲科;C~D.天牛科 stn1前胸腹板;stn2中胸腹板;stn3后胸腹板;cx1前足基节;cx2中足基节;cx3后足基节; sp前胸腹板刺;cxc基节窝 19.叶甲科Chrysomelidae(图47A~E) 体常有金属光泽;触角丝状或锯齿状,11节,少数9~10节,不及体长之半;复眼卵圆;跗节隐5节。幼虫蛃型。重要种类:黄曲条跳甲 Phyllotreta striolata (Fabricius)。 图47 叶甲科 A.头;B.跗节;C.幼虫;D.蛹;E.成虫 20.拟步甲科Tenebrionidae (图48D) 外形似步甲。前唇明显;鞘翅有发达假缘折;前足基节窝闭式;跗节式5-5-4。幼虫似金针虫,但腹末无金针虫那样的骨质突起和伪足。重要饲 料昆虫:黄粉虫Tenebrio molitor L.。 21.芫青科Meloidae(图48E) 头与前胸等宽;下口式;鞘翅较软;前足基节窝开放式;跗节式5-5-4;爪二分裂。幼虫复变态,捕食蝗虫卵等。药用种类:大斑芫青Mylabris phalerata (Pallas)。 图48 鞘翅目分科 A.瓢虫科成虫;B.瓢虫科蛹;C.瓢虫科幼虫;D,拟步甲科;E.芫青科 22.瓢虫科Coccinellidae(图48A~C) 体半球形;鞘翅有缘折;第1腹板上有后基线;跗节隐4节。多为肉食性,少数为植食性。肉食性瓢虫成虫体背无毛,有光泽,触角着生于两复眼 之前,上颚具基齿;幼虫行动活泼,体前端阔,后方狭,体上有软的肉刺及瘤粒。著名种类:澳洲瓢虫Rodolia cardinalis(Mulsant)。植食性瓢虫 成虫体被细毛,无光泽,触角着生于复眼之间,上颚不具基齿;幼虫爬动缓慢,体背多具硬而分叉状枝刺。重要种类:马铃薯瓢虫Epilachna vigintiomaculata Motschulsky。 23.三锥象甲科Brentidae(图49A) 触角9~11节,非膝状;喙细长,直,约与前胸等长;前足基节半圆形;后胸腹板约与腹部等长;可见腹板5节;跗节隐5节。常见种类:甘薯小 象甲Cylas formicarius (Fabricius)。 图49 鞘翅目分科 A.三锥象甲科;B.象甲科C.小蠹科 24.象甲科Curculionidae(图49B) 触角11节,膝状;喙弯曲;跗节隐5节。本科为鞘翅目最大科。重要种类:谷象Sitophilus granarius (L.)
25.小蠹科Scolytidae(图49C) 触角膝状,端部3~4节成锤状:头后部被前胸背板覆盖,胫节扁,具齿列:前胸长于体长的1/3。重要种类:落叶松小蠹 Scolytus (Ruguloscolytus)morawitzi Semenov. 作业与思考题 1.请简述鞘翅目与人类的关系。 2.如何区别植食性瓢虫与捕食性瓢虫? 3.鞘翅目的幼虫分属哪些类型?
25.小蠹科Scolytidae(图49C) 触角膝状,端部3~4节成锤状;头后部被前胸背板覆盖,胫节扁,具齿列;前胸长于体长的1/3。重要种类:落叶松小蠹 Scolytus (Ruguloscolytus) morawitzi Semenov。 作业与思考题 1.请简述鞘翅目与人类的关系。 2.如何区别植食性瓢虫与捕食性瓢虫? 3.鞘翅目的幼虫分属哪些类型?